Another theme, inextricably tied to the origin debate, was the investigation of the germ theory of disease. Edgar March Crookshank (1858-1928), a student of both Lister and the German pioneer Robert Koch, established England's first bacteriology laboratory. Alexander Ogston (1844-1929) of Aberdeen studied under the masters of Europe and, in 1882, discovered and named the genus Staphylococcus, a major pathogen. Preeminent among the medical bacteriologists, however, was Joseph Lister (1827-1912), who revolutionized surgery by his use of antisepsis.